From “Genetic Domains in Tall Bearded Iris Flowers” in Irises: The Bulletin of the American Iris Society, Volume 93 No. 2 (April 2012), edited by Kelly D. Norris:
“Rhizomes, leaves, stems, standards, falls, hafts, beards, styles, even veins in the iris flower have different functions โ hence the term functional domains. These domains have their own regulatory genes that form their respective organs during development. Some of these regulatory genes turn on or off the carotenoid and anthocyanin color pathways — the sequence of biochemical reactions necessary to produce a colored pigment….
“There are at least six genetic domains in tall bearded iris flowers. These are the standards, falls, hafts, beards, veins, and styles. Remember, the falls, standards, and styles differentiate from different whorls of developing plant cells and take their own color pathways with them. Therefore, these floral parts should not be considered as one unit. Each of these domains has a duplicate gene that regulates its color pathway. We know these genetic domains exist because of the different colors expressed….
“A solid colored flower, or self, has its color genes in sync โ the standards, falls, hafts, beards, veins, and styles all express the same alleles for color. Thus the duplicate regulatory genes in a self have similar alleles, but once the synchrony between alleles is changed a wonderful world of color combinations opens up….
“Iris breeding for the past 100 years has focused on changing the synchrony between floral genetic domains to develop new color combinations (not new pigments). For example, first there was the amoena and then came the reverse amoena. The development of amoena, bitone, and luminata flowers has been a progression of changing the synchrony between duplicate genes in different domains.”
From “In Zurich” in So That It Flower: A Gathering of Poems by Melville Cane:
In Zurich,
Au lac,
As soon as it is June,
They deck the tables on the terraces
With yellow irises,
And serve a brand of scrupulously fine
Gay sunshine,
To mix with equally clear and sunny
Amber honey
And butter-balls, like curled tea-roses.
Then, if you so incline,
A draught, or two, or three, of wine;
And, as the blood begins
To sing with the violins,
One muses
And drowses.
Hello!
This is the second of two posts with photographs of irises with yellow standards and white falls that I took at Oakland Cemetery earlier this year. In the first post — Yellow and White Irises: Reverse Amoena Patterns (1 of 2) — I included irises planted in some of Oakland’s oldest landscapes where the irises hung out in solitary groups. In this post, we see some rejuvenated landscapes (like the first gallery image below) where the irises have complementary neighbors: the tall white-flowered Veronica spicata (also known as Spike Speedwell); and purple-flowered Verbena bonariensis (also known as Tall Verbena, Brazilian Verbena, or Purpletop Vervain), which I’ve positioned as the subject of a few of the photos with the irises in a supporting role. Is it just me, or do “Spike Speedwell” and “Purpletop Vervain” sound like they could be the names of cartoon characters?
With this post, I’ve reached the end of spring and summer 2026 photographs from Oakland. In past years, I’ve photographed lots of lilies in July and August, but doing so has meant that I’m still working on lily photos into fall and early winter, leading me to skip or miss out on photographing autumn color with the attention it deserves. Given how many asters and mums Oakland produces after summer subsides — and the variety of colors that its trees and shrubs produce — I hope to spend more time in an autumn frame of mind before the holidays roll around, without the lily photos pressuring me (!!) to edit, post, and write about them.
This, of course, assumes the temperatures start to turn cool here soon — we’ve been in the mid-to high-nineties every day throughout August and September so far — as asters, mums, and all the shrubs and trees really do use the cooling temperatures to tell them how to proceed toward fall. There’s no evidence of that cooling yet…

… but if we start dropping into the 50s at night this weekend and next week…

… the plants will likely take notice. And I will too.
Having lived half my life in northern New York, I have lingering nostalgia for the temperature changes that occur there between August and September, right when summer’s humid heat has worn out its welcome. Memories of my earliest visits to Oakland add to that nostalgia: I first explored its history and architecture extensively in October and November of 2007 for a series of research papers. So I’m planning regular stomps through the grounds of Oakland (as well as nearby Grant Park) in jeans, sneaks, and a nice fluffy jacket this fall — and recently bought a more landscape-appropriate new (used) camera lens just for that purpose.
The excerpt at the top of this post describes six “genetic domains” for tall bearded irises like these reverse amoenas, explaining how the flower’s standards, falls, hafts, beards, veins, and styles are all considered distinct (and open to manipulation) by geneticists producing iris variants. While I don’t claim to understand that part of the science well enough to explain it (yet), I still thought it was interesting that these genetic domains can be useful to a photographer (or any observer) as a way of isolating and focusing on various color regions our cameras capture. To help me visualize that better, I asked Claude.AI to produce a drawing mapping the genetic domains to their locations on the flower.
Using photos like this as guidance…

… Claude came up with this, a skeleton version of the reverse amoena irises that includes the domains mentioned in the excerpt (except veins, which could appear anywhere among the other sections).

If we put them side-by-side, it’s easy to see how the drawing is an abstract map to the living color regions of the flower I photographed….


… and it’s these regions, along with the color and texture transitions between them, that can be the most compelling ones to zoom in on. Close-up and macro photos of irises, in particular, are often composed to fill the frame with the regional transitions — not necessarily because the photographer knew about genetic domains and their color influences, but because these flower parts so deftly illustrate the ability of an iris (through genetic manipulation or natural mutations) to produce distinct arrangements of color, form, and texture within and at the boundaries of these regions — and color, form, and texture are so consequential to photography, photographic theory, and other kinds of image production.
Thanks for reading and taking a look!








































































































